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So, there are no reestriction about how direction that you can take, for example, you can take some traits nodes from Elamintes and after that maybe you will like to go to Egypt old-kingdom tree, as long as you accomplished the requirements of each tree -if exist-, in general terms always you can progress through trees that are reasonably closely, your faction can change passively his acclimatation based on where is placed over the world map, happens more faster at early game meanwhile you are nomad but still can change at midle or late game if you want to do.
To unlock new nodes from a particular tier requires have unlocked the same nodes than that tier row less one (simple formula: tier row -1) , that is like if you want to unlock nodes from tier 2 you need at least to have one node, or if you want to unlock any node from tier 5 you need to have at least 4 nodes of that branch.
Currently some civilization and cultures like Rome Republic have up to 5 branchs specializations, each branch can contain up to 5 tiers so that means up to 15 nodes per each branch, also each tree include tier bonuses if you unlock enough nodes like from picture of early indoeuropeans desing, that in this case unlocks an architectonic collection.
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