Photosynthesis
Alternative methods of carbon
fixation in hot, arid climates
Your goals
◊ Explain how C4 and CAM plants have
adapted to live in hot, windy climates
You may want to set up your
notes like this
C3 Plants C4 Plants CAM Plants
Plant evolutionary history
Life began in water
The first plants were aquatic
Since plants evolved to living out of water,
they have had a hard time dealing with
dehydration
Plant adaptations
Most adaptations involve some sort of
trade off
For plants, the trade off with
photosynthesis is that they lose water to
the environment through transpiration
A little plant anatomy
The openings on leaves are called stomataThe openings on leaves are called stomata
This is where CO2 and O2 can enter and exit
the plant
Water is also lost through the stomata
Plants often close the stomata on very hotPlants often close the stomata on very hot
days, but COdays, but CO22 intake is cut offintake is cut off
C3 plants
Most plants
Fix carbon byFix carbon by
attaching COattaching CO22 to RuBPto RuBP
“3” comes from the 3-carbonmolecule produced after CO2is fixed
C3 Plants
On hot days, they close their stomata
part way
Produce little sugar then
C3 plants
As the COAs the CO22 that the plant does havethat the plant does have
gets used up, rubisco will fix Ogets used up, rubisco will fix O22 insteadinstead
and send it into the Calvin cycle instead
of CO2
Peroxisomes & mitochondria split the
new compound and release CO2
This is called photorespiration
Occurs in the
light
Consumes oxygen and
releases CO2
Photorespiration
uses ATP but
makes no ATP
Photorespiration
makes no sugars
PhotorespirationPhotorespiration
decreasesdecreases
photosynthesisphotosynthesis
outputoutput
So why would any
plant do this…
Photorespiration mayPhotorespiration may
be an evolutionarybe an evolutionary
leftoverleftover
In early atmosphere,
with little O2, it didn’t
matter if Rubisco had an
affinity for O2
Today, with so much O2
in the atmosphere, it is
inevitable that some O2
will be fixed instead of
C3 Crop plants
Rice, wheat, soybeansRice, wheat, soybeans
May lose as much as 50% of the
Carbon fixed in the Calvin Cycle
to photorespiration
3 Things
What are 3 things you know for sure
about C3 plants?
C4 plants
C4 plants have
adaptations that
allow them to
minimize the
effects of
photorespiration
Many are
tropical grasses
C4 plants
Have an alternate
means of fixing
carbon
C4 plants fix carbon
out in the
cytoplasm before it
enters the Calvin
Cycle
“4” comes from the 4-carboncompound that is formed when CO2
is fixed in C4 plants
C4 plant adaptations
1st step is binding CO2 to PEP
(phosphoenolpyruvate) by the enzyme PEP
carboxylase to make a 4-C compound:
oxaloacetate
PEP has a high affinity for CO2 and none for
O2
2. The 4-carbon compound enters
photosynthetic cellsphotosynthetic cells
CO2 is released from PEP and the Calvin
cycle continues as normal
This adaptation allows C4 plants to keep a
high concentration of CO2 in the
photosynthetic cells, preventing Rubisco from
binding to O2 instead of CO2
C4 plants thrive in hot
climates where
stomata will be
closed often
Some important C4
plants are sugarcane,
corn, tropical grasses
3 Things
What are 3 things you know for sure
about C4 plants?
CAM plants
Another alternate method of carbon
fixation
Common in succulents like cacti
CAM plants
CAM plants keep stomata closed during
the day to minimize water loss
Only open stomata at night
Closed stomata during the day
means no CO2 entering the
plant during the day
CAM plants
When stomata are
open at night, CAM
plants take in CO2 and
fix it to PEP
This is called
crassulacean acid
metabolism
Crassulaceae is the
name of the family ofplants that include
succulents
CAM plants
Cells of CAM plants
store the organic
molecules that CO2 was
fixed to in their
vacuoles until morning
Light reactions then
produce ATP and
NADPH to drive the
Calvin Cycle
The CO2 in the organic
molecules is released
for use in the Calvin
Cycle
CAM plants
Important CAM plants include
pineapple and agave
3 Things
What are 3 things you know for sure
about CAM plants?
Comparison of C3, C4, and CAM plants
C3 plants C4 plants CAM plants
Most plants
Tropical grasses
like corn,
sugarcane
Succulents,
pineapple, agave
Fix carbon in Calvin
cycle - attach CO2
to RuBP
Fix carbon in
cytoplasm - attach
CO2 to PEP
Fix carbon at night
only, fix it to
organic molecules
Enzyme - Rubisco Enzyme – PEP-ase Enzyme – PEP-ase
Most energy
efficient method
1/2 way between
these two
Best water
conservation
Loses water
through
photorespiration
Loses less water Loses least water

AP Biology Photosynthesis C3 C4 and CAM plants

  • 1.
    Photosynthesis Alternative methods ofcarbon fixation in hot, arid climates
  • 2.
    Your goals ◊ Explainhow C4 and CAM plants have adapted to live in hot, windy climates
  • 3.
    You may wantto set up your notes like this C3 Plants C4 Plants CAM Plants
  • 4.
    Plant evolutionary history Lifebegan in water The first plants were aquatic Since plants evolved to living out of water, they have had a hard time dealing with dehydration
  • 5.
    Plant adaptations Most adaptationsinvolve some sort of trade off For plants, the trade off with photosynthesis is that they lose water to the environment through transpiration
  • 6.
    A little plantanatomy The openings on leaves are called stomataThe openings on leaves are called stomata This is where CO2 and O2 can enter and exit the plant Water is also lost through the stomata Plants often close the stomata on very hotPlants often close the stomata on very hot days, but COdays, but CO22 intake is cut offintake is cut off
  • 7.
    C3 plants Most plants Fixcarbon byFix carbon by attaching COattaching CO22 to RuBPto RuBP “3” comes from the 3-carbonmolecule produced after CO2is fixed
  • 8.
    C3 Plants On hotdays, they close their stomata part way Produce little sugar then
  • 9.
    C3 plants As theCOAs the CO22 that the plant does havethat the plant does have gets used up, rubisco will fix Ogets used up, rubisco will fix O22 insteadinstead and send it into the Calvin cycle instead of CO2
  • 10.
    Peroxisomes & mitochondriasplit the new compound and release CO2 This is called photorespiration Occurs in the light Consumes oxygen and releases CO2
  • 11.
    Photorespiration uses ATP but makesno ATP Photorespiration makes no sugars PhotorespirationPhotorespiration decreasesdecreases photosynthesisphotosynthesis outputoutput
  • 12.
    So why wouldany plant do this… Photorespiration mayPhotorespiration may be an evolutionarybe an evolutionary leftoverleftover In early atmosphere, with little O2, it didn’t matter if Rubisco had an affinity for O2 Today, with so much O2 in the atmosphere, it is inevitable that some O2 will be fixed instead of
  • 13.
    C3 Crop plants Rice,wheat, soybeansRice, wheat, soybeans May lose as much as 50% of the Carbon fixed in the Calvin Cycle to photorespiration
  • 14.
    3 Things What are3 things you know for sure about C3 plants?
  • 15.
    C4 plants C4 plantshave adaptations that allow them to minimize the effects of photorespiration Many are tropical grasses
  • 16.
    C4 plants Have analternate means of fixing carbon C4 plants fix carbon out in the cytoplasm before it enters the Calvin Cycle “4” comes from the 4-carboncompound that is formed when CO2 is fixed in C4 plants
  • 17.
    C4 plant adaptations 1ststep is binding CO2 to PEP (phosphoenolpyruvate) by the enzyme PEP carboxylase to make a 4-C compound: oxaloacetate PEP has a high affinity for CO2 and none for O2
  • 18.
    2. The 4-carboncompound enters photosynthetic cellsphotosynthetic cells CO2 is released from PEP and the Calvin cycle continues as normal This adaptation allows C4 plants to keep a high concentration of CO2 in the photosynthetic cells, preventing Rubisco from binding to O2 instead of CO2
  • 19.
    C4 plants thrivein hot climates where stomata will be closed often Some important C4 plants are sugarcane, corn, tropical grasses
  • 20.
    3 Things What are3 things you know for sure about C4 plants?
  • 21.
    CAM plants Another alternatemethod of carbon fixation Common in succulents like cacti
  • 22.
    CAM plants CAM plantskeep stomata closed during the day to minimize water loss Only open stomata at night
  • 23.
    Closed stomata duringthe day means no CO2 entering the plant during the day
  • 24.
    CAM plants When stomataare open at night, CAM plants take in CO2 and fix it to PEP This is called crassulacean acid metabolism Crassulaceae is the name of the family ofplants that include succulents
  • 25.
    CAM plants Cells ofCAM plants store the organic molecules that CO2 was fixed to in their vacuoles until morning Light reactions then produce ATP and NADPH to drive the Calvin Cycle The CO2 in the organic molecules is released for use in the Calvin Cycle
  • 26.
    CAM plants Important CAMplants include pineapple and agave
  • 27.
    3 Things What are3 things you know for sure about CAM plants?
  • 29.
    Comparison of C3,C4, and CAM plants C3 plants C4 plants CAM plants Most plants Tropical grasses like corn, sugarcane Succulents, pineapple, agave Fix carbon in Calvin cycle - attach CO2 to RuBP Fix carbon in cytoplasm - attach CO2 to PEP Fix carbon at night only, fix it to organic molecules Enzyme - Rubisco Enzyme – PEP-ase Enzyme – PEP-ase Most energy efficient method 1/2 way between these two Best water conservation Loses water through photorespiration Loses less water Loses least water