Unit 8-Evolution
15.1 Darwin’s Theory of Natural Selection
15.2 Evidence of Evolution
15.3 Shaping Evolutionary Theory
Name _________________________ Period _______
15.1 Darwin’s Theory of Natural Selection
• Main idea:
• Objectives
– Discuss the evidence that convinced Darwin that
species could change over time.
– List the four principles of natural selection.
– Show how natural selection could change a
population
• New Vocabulary
– Selective breeding, Artificial selection, Natural selection,
Evolution
Developing the Theory of Natural
Selection
• Charles Darwin boarded the
HMS Beagle in 1831.
• Most believed the world was
about 6,000 years old.
• During the ship’s five year
voyage, Darwin made
extensive collections of
rocks, fossils, plants and
animals.
• He also read Charles Lyell’s Principles of
Geology – a book proposing the Earth was
millions of years old.
Developing the Theory of Natural
Selection
• In 1835, the Beagle arrived in the Galapagos
Islands off the coast of South America.
• Darwin began to collect mockingbirds, finches,
and other animals on the four islands.
• He noticed that the different islands seemed
to have their own, slightly different varieties
of animals.
HMS Beagle
Developing the Theory of Natural
Selection
• Almost every
specimen that
Darwin had
collected on
the islands was
new to
European
scientists.
• Darwin suspected populations from the mainland
changed after reaching the Galápagos.
Galápagos Islands
Developing the Theory of Natural
Selection
• Darwin hypothesized that new species could
appear gradually through small changes in
ancestral species.
• Darwin inferred that if humans could change
species by artificial selection (selective
breeding), then perhaps the same process
could work in nature.
Developing the Theory of Natural
Selection
• Darwin called his theory Natural Selection
Natural SelectionNatural Selection
• He reasoned that, given
enough time, natural
selection could modify a
population enough to
produce a new species.
Basic Principles of Natural Selection
The Origin of Species
• Darwin published On the Origin of Species by
Means of Natural Selection in 1859.
• Today, biologists use the term evolution to
define changes in groups of organisms
through time.
• Darwin’s theory of natural selection is NOT
the same as evolution. Natural selection is a
means of explaining HOW evolution works.
15.2 Evidence of Evolution
• Main idea:
• Objectives:
– Describe how fossils provide evidence of evolution.
– Discuss morphological evidence of evolution.
– Explain how physiology and biochemistry provide
evidence of evolution.
• New Vocabulary

Derived trait

Ancestral trait

Homologous structure

Vestigial structure

Analogous structure

Embryo

Fitness

Mimicry

Camouflage
Support for Evolution
• The theory of evolution states that all organisms
on Earth have descended from a common
ancestor.
• The fossil record offers some of the most
significant evidence of evolutionary change.
– Fossils provide a record of species that lived
long ago.
– Fossils show that ancient species share
similarities with species that now live on Earth.
• The giant armadillo-like
glyptodont, Glyptodon,is an
extinct animal that Darwin
thought must be related to
the living armadillos of
today.
• The fossil record provides
information for determining
the ancestry of organisms
and the patterns of
evolution.
Support for Evolution
The Fossil Record
Support for Evolution
Two major classes of traits:
•Derived traits are newly evolved features,
such as feathers, that do not appear in the
fossils of common ancestors.
•Ancestral traits are more primitive
features, such as teeth and tails, that do
appear in ancestral forms.
The Fossil Record
Support for Evolution
• Transitional fossils provide detailed patterns
of evolutionary change for ancestors of many
modern animals, including mollusks, horses,
whales and humans.
Example:
•Archeopteryx

Shares features of
both dinosaurs and
birds.
The Fossil Record
Support for Evolution
• Homologous
structures are
anatomically
similar structures
inherited from a
common ancestor.
• The forelimbs of
vertebrates are
adapted for
different uses, but
they all have
similar bones.
• Similar structure; different
function.
Comparative Anatomy
Support for Evolution
• Vestigial structures are
the reduced forms of
functional structures in
other organisms.
• Evolutionary theory
predicts that features of
ancestors that no longer
have a function for that
species will become
smaller over time until
they are lost.• Examples: Snake pelvis and
human appendix.
Comparative Anatomy
Support for Evolution
• Analogous structures can be
used for the same purpose and
can be similar in construction,
but are not inherited from a
common ancestor.
• Show that functionally similar
features can evolve
independently in similar
environments.
• Different structure; same
function;
Comparative Anatomy
Support for Evolution
• An embryo is an early pre-birth stage of an
organism’s development.
• Vertebrate
embryos exhibit
homologous
structures during
certain phases of
development but
become totally
different
structures in the
adult forms.
Comparative Embryology
Comparative Biochemistry
• Common
ancestry can
be seen in the
complex
metabolic
molecules that
many different
organisms
share.
Support for Evolution
Geographic Distribution
• The distribution of plants and animals that
Darwin saw during his travels first suggested
evolution to Darwin.
Rabbit in Europe Mara in S. America
Support for Evolution
• Scientists have confirmed
and expanded Darwin’s
study of the distribution
of plants and animals
around the world in a
field of study now called
biogeography.
• Evolution is intimately linked with
climate and geological forces.
Geographic Distribution
Support for Evolution
Adaptation
• An adaptation is a trait
shaped by natural selection
that increases an
organism’s reproductive
success.
• Fitness is a measure of the relative contribution
an individual trait makes to the next generation.
It is often measured as the number of
reproductively viable offspring that an organism
produces in the next generation.
• Camouflage allows organisms
to become almost invisible to
predators. Some species have
evolved morphological
adaptations that allow them
to blend in with their
environments.
• Mimicry is another
morphological adaptation
that allows one species to
evolve to resemble another
species.
Adaptation
Types of Adaptation
• Antimicrobial resistance
- An antibiotic is a
medicine that slows or
kills the growth of
bacteria.
– Some bacteria have
evolved a resistance
to certain antibiotics.

People infected with resistant bacteria can never get rid of it.
Adaptation
Types of Adaptation
15.3 Shaping Evolutionary Theory
• Main idea:
• Objectives:
– Discuss patterns observed in evolution.
– Describe factors that influence speciation.
– Compare gradualism with punctuated equilibrium.
• Review Vocabulary
– Allele: alternative forms of a character trait that can be
inherited.
15.3 Shaping Evolutionary Theory
• Hardy-Weinberg Principle
• Genetic drift
• Founder effect
• Bottleneck
• Punctuated Equilibrium
• Sexual selection
New Vocabulary
Mechanisms of Evolution
• Darwin’s theory of natural selection
remains the central theme of evolution
• Scientists of today know that natural
selection is not the only mechanism of
evolution
• Evolution occurs at the population level,
with genes as the raw material.
Mechanisms of Evolution
• Hardy-Weinberg Principle states that when allelic
frequencies remain constant, a population is in
genetic equilibrium.
Population Genetics

Even though the number of owls doubled, the ratio of gray
to red owls remained the same.
Mechanisms of Evolution
• p2
+ 2pq + q2
= 1
• This equation allows us to determine the
equilibrium frequency of each genotype in the
population:
• homozygous dominant (p2
)
• heterozygous (2pq)
• homozygous recessive (q2
)
– Note the sum of these frequencies equals one.
Population Genetics
Mechanisms of Evolution
A population in genetic equilibrium must meet all five
conditions
•The population is very large
•No immigration or emigration
•Mating is random
•Mutations do not occur (no new variations)
•Natural selection does not occur
Population Genetics
• Genetic Drift is a change in the allelic
frequencies in a population that is due to
chance.
• In smaller populations, the effects of genetic
drift becomes more pronounced, and the
chance of losing an allele becomes greater.
• Examples: Founder Effect & Bottleneck
Mechanisms of Evolution
Genetic Drift
• Founder Effect occurs when a small sample of a
population settles in a location separated from the
rest of the population
Genetic Drift
• Alleles that were
uncommon in the
original population
might be common in
the new population.
Mechanisms of Evolution
• Alleles that were uncommon in the original
population might be common in the new population.
• Bottleneck occurs
when a population
declines to a very
low number and
then rebounds
Mechanisms of Evolution
Genetic Drift

Can decrease
genetic variety.
• A population in genetic equilibrium
experiences no gene flow, however, few
populations are isolated
• Random movement of individuals between
populations reduces differences between
populations
Mechanisms of Evolution
Gene Flow
• Rarely is mating completely
random in a population.
• Usually individuals mate
with individuals in close
proximity.
Nonrandom mating
Mechanisms of Evolution
• This promotes inbreeding and could lead to a change
in allelic proportions favoring individuals that are
homozygous for particular traits.
• A mutation is a random
change in genetic material.
• Occasionally mutations
provide an advantage to
organisms and become more
common in subsequent
generations.
• Basis upon which natural
selection works.
Mechanisms of Evolution
Mutation
Select the individuals that are best
adapted for survival and
reproduction
•Stabilizing selection
– eliminate extreme
expressions
• Birth Weights
•Directional selection
– makes organisms more fit.
• Peppered Moths
•Disruptive selection
– process that splits a
population into two groups.
• Lake Erie Water Snakes
Mechanisms of Evolution
Natural Selection
• Sexual selection operates in
populations where males and
females differ significantly in
appearance.
– Males are largest and most colorful
in the group
– More likely target for predators yet
bodies enhance reproductive
success
• Examples: Peacocks & Deer
Natural Selection
Mechanisms of Evolution
• prevents reproduction
by making fertilization
unlikely.
– Eastern and Western
meadowlarks similar in
appearance but their
mating songs separate
them behaviorally.
Reproductive Isolation
Prezygotic Isolation
• Occurs when
fertilization has
occurred but a
hybrid offspring
cannot develop or
reproduce.
Reproductive Isolation
Postzygotic Isolation

A Liger is the offspring of a male lion and
a female tiger. Ligers are sterile.
• A physical barrier divides one population into
two or more populations.
Speciation
Allopatric speciation
• For speciation to occur, a population must
diverge and then be reproductively isolated.
• Species evolves into a new species without a
physical barrier
Speciation
Sympatric speciation

Apple maggot flies lay
eggs on apples or
hawthorns depending
on the type of fruit the
grew up on.
• For speciation to occur, a population must
diverge and then be reproductively isolated.
• Can occur in a relatively short time when one
species gives rise to many different species in
response to the creation of new habitat or some
other ecological opportunity.
Patterns of Evolution
Adaptive Radiation (aka divergent evolution)
• Follows large-
scale
extinction
events

Cichlid fish of
Lake Victoria
in Africa.
• The relationship between two species might be so
close that the evolution of one species affects the
evolution of the other species.
• Mutualism – occurs when two species benefit each
other.
Coevolution
Patterns of Evolution
• The plant evolves a chemical
defense
• The insects evolve the
biochemistry to resist the
defense
• Response continues to escalate.
Patterns of Evolution
Coevolution
• Coevolutionary arms race – one species can
evolve a parasitic dependency on another
species.
• A plant and an insect that is dependent on the
plant for food
• Unrelated species
evolve similar traits
even though they live in
different parts of the
world.
• Occurs in environments
that are geographically
far apart but have
similar ecology and
climate.
Patterns of Evolution
Convergent Evolution
• Punctuated
equilibrium
explains rapid
spurts of genetic
change causing
species to diverge
quickly.
Rate of Speciation
Patterns of Evolution
• Evolution proceeds in small, gradual steps =
gradualism.

Chapter 15 Evolution - All Sections 15.1 - 15.3

  • 1.
    Unit 8-Evolution 15.1 Darwin’sTheory of Natural Selection 15.2 Evidence of Evolution 15.3 Shaping Evolutionary Theory Name _________________________ Period _______
  • 2.
    15.1 Darwin’s Theoryof Natural Selection • Main idea: • Objectives – Discuss the evidence that convinced Darwin that species could change over time. – List the four principles of natural selection. – Show how natural selection could change a population • New Vocabulary – Selective breeding, Artificial selection, Natural selection, Evolution
  • 3.
    Developing the Theoryof Natural Selection • Charles Darwin boarded the HMS Beagle in 1831. • Most believed the world was about 6,000 years old. • During the ship’s five year voyage, Darwin made extensive collections of rocks, fossils, plants and animals. • He also read Charles Lyell’s Principles of Geology – a book proposing the Earth was millions of years old.
  • 4.
    Developing the Theoryof Natural Selection • In 1835, the Beagle arrived in the Galapagos Islands off the coast of South America. • Darwin began to collect mockingbirds, finches, and other animals on the four islands. • He noticed that the different islands seemed to have their own, slightly different varieties of animals. HMS Beagle
  • 5.
    Developing the Theoryof Natural Selection • Almost every specimen that Darwin had collected on the islands was new to European scientists. • Darwin suspected populations from the mainland changed after reaching the Galápagos. Galápagos Islands
  • 6.
    Developing the Theoryof Natural Selection • Darwin hypothesized that new species could appear gradually through small changes in ancestral species. • Darwin inferred that if humans could change species by artificial selection (selective breeding), then perhaps the same process could work in nature.
  • 7.
    Developing the Theoryof Natural Selection • Darwin called his theory Natural Selection Natural SelectionNatural Selection • He reasoned that, given enough time, natural selection could modify a population enough to produce a new species.
  • 8.
    Basic Principles ofNatural Selection
  • 9.
    The Origin ofSpecies • Darwin published On the Origin of Species by Means of Natural Selection in 1859. • Today, biologists use the term evolution to define changes in groups of organisms through time. • Darwin’s theory of natural selection is NOT the same as evolution. Natural selection is a means of explaining HOW evolution works.
  • 10.
    15.2 Evidence ofEvolution • Main idea: • Objectives: – Describe how fossils provide evidence of evolution. – Discuss morphological evidence of evolution. – Explain how physiology and biochemistry provide evidence of evolution. • New Vocabulary  Derived trait  Ancestral trait  Homologous structure  Vestigial structure  Analogous structure  Embryo  Fitness  Mimicry  Camouflage
  • 11.
    Support for Evolution •The theory of evolution states that all organisms on Earth have descended from a common ancestor. • The fossil record offers some of the most significant evidence of evolutionary change. – Fossils provide a record of species that lived long ago. – Fossils show that ancient species share similarities with species that now live on Earth.
  • 12.
    • The giantarmadillo-like glyptodont, Glyptodon,is an extinct animal that Darwin thought must be related to the living armadillos of today. • The fossil record provides information for determining the ancestry of organisms and the patterns of evolution. Support for Evolution The Fossil Record
  • 13.
    Support for Evolution Twomajor classes of traits: •Derived traits are newly evolved features, such as feathers, that do not appear in the fossils of common ancestors. •Ancestral traits are more primitive features, such as teeth and tails, that do appear in ancestral forms. The Fossil Record
  • 14.
    Support for Evolution •Transitional fossils provide detailed patterns of evolutionary change for ancestors of many modern animals, including mollusks, horses, whales and humans. Example: •Archeopteryx  Shares features of both dinosaurs and birds. The Fossil Record
  • 15.
    Support for Evolution •Homologous structures are anatomically similar structures inherited from a common ancestor. • The forelimbs of vertebrates are adapted for different uses, but they all have similar bones. • Similar structure; different function. Comparative Anatomy
  • 16.
    Support for Evolution •Vestigial structures are the reduced forms of functional structures in other organisms. • Evolutionary theory predicts that features of ancestors that no longer have a function for that species will become smaller over time until they are lost.• Examples: Snake pelvis and human appendix. Comparative Anatomy
  • 17.
    Support for Evolution •Analogous structures can be used for the same purpose and can be similar in construction, but are not inherited from a common ancestor. • Show that functionally similar features can evolve independently in similar environments. • Different structure; same function; Comparative Anatomy
  • 18.
    Support for Evolution •An embryo is an early pre-birth stage of an organism’s development. • Vertebrate embryos exhibit homologous structures during certain phases of development but become totally different structures in the adult forms. Comparative Embryology
  • 19.
    Comparative Biochemistry • Common ancestrycan be seen in the complex metabolic molecules that many different organisms share. Support for Evolution
  • 20.
    Geographic Distribution • Thedistribution of plants and animals that Darwin saw during his travels first suggested evolution to Darwin. Rabbit in Europe Mara in S. America Support for Evolution
  • 21.
    • Scientists haveconfirmed and expanded Darwin’s study of the distribution of plants and animals around the world in a field of study now called biogeography. • Evolution is intimately linked with climate and geological forces. Geographic Distribution Support for Evolution
  • 22.
    Adaptation • An adaptationis a trait shaped by natural selection that increases an organism’s reproductive success. • Fitness is a measure of the relative contribution an individual trait makes to the next generation. It is often measured as the number of reproductively viable offspring that an organism produces in the next generation.
  • 23.
    • Camouflage allowsorganisms to become almost invisible to predators. Some species have evolved morphological adaptations that allow them to blend in with their environments. • Mimicry is another morphological adaptation that allows one species to evolve to resemble another species. Adaptation Types of Adaptation
  • 24.
    • Antimicrobial resistance -An antibiotic is a medicine that slows or kills the growth of bacteria. – Some bacteria have evolved a resistance to certain antibiotics.  People infected with resistant bacteria can never get rid of it. Adaptation Types of Adaptation
  • 25.
    15.3 Shaping EvolutionaryTheory • Main idea: • Objectives: – Discuss patterns observed in evolution. – Describe factors that influence speciation. – Compare gradualism with punctuated equilibrium. • Review Vocabulary – Allele: alternative forms of a character trait that can be inherited.
  • 26.
    15.3 Shaping EvolutionaryTheory • Hardy-Weinberg Principle • Genetic drift • Founder effect • Bottleneck • Punctuated Equilibrium • Sexual selection New Vocabulary
  • 27.
    Mechanisms of Evolution •Darwin’s theory of natural selection remains the central theme of evolution • Scientists of today know that natural selection is not the only mechanism of evolution • Evolution occurs at the population level, with genes as the raw material.
  • 28.
    Mechanisms of Evolution •Hardy-Weinberg Principle states that when allelic frequencies remain constant, a population is in genetic equilibrium. Population Genetics  Even though the number of owls doubled, the ratio of gray to red owls remained the same.
  • 29.
    Mechanisms of Evolution •p2 + 2pq + q2 = 1 • This equation allows us to determine the equilibrium frequency of each genotype in the population: • homozygous dominant (p2 ) • heterozygous (2pq) • homozygous recessive (q2 ) – Note the sum of these frequencies equals one. Population Genetics
  • 30.
    Mechanisms of Evolution Apopulation in genetic equilibrium must meet all five conditions •The population is very large •No immigration or emigration •Mating is random •Mutations do not occur (no new variations) •Natural selection does not occur Population Genetics
  • 31.
    • Genetic Driftis a change in the allelic frequencies in a population that is due to chance. • In smaller populations, the effects of genetic drift becomes more pronounced, and the chance of losing an allele becomes greater. • Examples: Founder Effect & Bottleneck Mechanisms of Evolution Genetic Drift
  • 32.
    • Founder Effectoccurs when a small sample of a population settles in a location separated from the rest of the population Genetic Drift • Alleles that were uncommon in the original population might be common in the new population. Mechanisms of Evolution • Alleles that were uncommon in the original population might be common in the new population.
  • 33.
    • Bottleneck occurs whena population declines to a very low number and then rebounds Mechanisms of Evolution Genetic Drift  Can decrease genetic variety.
  • 34.
    • A populationin genetic equilibrium experiences no gene flow, however, few populations are isolated • Random movement of individuals between populations reduces differences between populations Mechanisms of Evolution Gene Flow
  • 35.
    • Rarely ismating completely random in a population. • Usually individuals mate with individuals in close proximity. Nonrandom mating Mechanisms of Evolution • This promotes inbreeding and could lead to a change in allelic proportions favoring individuals that are homozygous for particular traits.
  • 36.
    • A mutationis a random change in genetic material. • Occasionally mutations provide an advantage to organisms and become more common in subsequent generations. • Basis upon which natural selection works. Mechanisms of Evolution Mutation
  • 37.
    Select the individualsthat are best adapted for survival and reproduction •Stabilizing selection – eliminate extreme expressions • Birth Weights •Directional selection – makes organisms more fit. • Peppered Moths •Disruptive selection – process that splits a population into two groups. • Lake Erie Water Snakes Mechanisms of Evolution Natural Selection
  • 38.
    • Sexual selectionoperates in populations where males and females differ significantly in appearance. – Males are largest and most colorful in the group – More likely target for predators yet bodies enhance reproductive success • Examples: Peacocks & Deer Natural Selection Mechanisms of Evolution
  • 39.
    • prevents reproduction bymaking fertilization unlikely. – Eastern and Western meadowlarks similar in appearance but their mating songs separate them behaviorally. Reproductive Isolation Prezygotic Isolation
  • 40.
    • Occurs when fertilizationhas occurred but a hybrid offspring cannot develop or reproduce. Reproductive Isolation Postzygotic Isolation  A Liger is the offspring of a male lion and a female tiger. Ligers are sterile.
  • 41.
    • A physicalbarrier divides one population into two or more populations. Speciation Allopatric speciation • For speciation to occur, a population must diverge and then be reproductively isolated.
  • 42.
    • Species evolvesinto a new species without a physical barrier Speciation Sympatric speciation  Apple maggot flies lay eggs on apples or hawthorns depending on the type of fruit the grew up on. • For speciation to occur, a population must diverge and then be reproductively isolated.
  • 43.
    • Can occurin a relatively short time when one species gives rise to many different species in response to the creation of new habitat or some other ecological opportunity. Patterns of Evolution Adaptive Radiation (aka divergent evolution) • Follows large- scale extinction events  Cichlid fish of Lake Victoria in Africa.
  • 44.
    • The relationshipbetween two species might be so close that the evolution of one species affects the evolution of the other species. • Mutualism – occurs when two species benefit each other. Coevolution Patterns of Evolution
  • 45.
    • The plantevolves a chemical defense • The insects evolve the biochemistry to resist the defense • Response continues to escalate. Patterns of Evolution Coevolution • Coevolutionary arms race – one species can evolve a parasitic dependency on another species. • A plant and an insect that is dependent on the plant for food
  • 46.
    • Unrelated species evolvesimilar traits even though they live in different parts of the world. • Occurs in environments that are geographically far apart but have similar ecology and climate. Patterns of Evolution Convergent Evolution
  • 47.
    • Punctuated equilibrium explains rapid spurtsof genetic change causing species to diverge quickly. Rate of Speciation Patterns of Evolution • Evolution proceeds in small, gradual steps = gradualism.