The document summarizes the characteristics of the phylum Chordata. It describes that chordates are defined by having a notochord, dorsal nerve chord, pharyngeal gill slits, post-anal tail, and an endostyle. The phylum is divided into two subphyla - Protochordata and Vertebrata. Protochordata includes hemichordates, calcichordates, urochordates, and cephalochordates. Urochordates are commonly called sea squirts and are mostly sessile, while cephalochordates include lancelets which have characteristics more developed than other protochordates. The phylum exhibits advancement over non-chordates in features
CHORDATA
Most advancedphylum
Have around 48100 species
Humans are intelligent & dominating animal of chordata
Phylum chordata created by Balfour (1880)
Chorda means Cord; refers to presence of a stiff rod like structure called “Notochord”
Notes means Back ;Chorda means Cord
3.
Animal belonging toPhylum Chordata are
fundamentally characterized by:
A notochord
A dorsal hollow nerve chord
Paired pharyngeal gill slits
Post anal tail
Presence of endostyle
4.
Advancement of Chordataover Non-
Chordata
Living endoskeleton which grow with the body
More efficient respiratory organs (Gills/lungs)
More efficient circulatory system with well developed
heart having cardiac muscles
Direction of blood flow first flows forward ventrally,then
dorsally and backwords
Better sense organs and tubular nervous system
Endocrine glands for efficient chemical coordination
5.
All Chordata are
Triploblastic
Only two paired appendages
Bilateral symmetry
Deuterostomes
Coelomate
Organ system level of organization
Segmented body
Presence of Endostyle (Thyroid gland)
Ventral contractile heart
6.
Classification of Chordates
Thephylum Chordata is divided in to two subphyla :-
Protochordata/Acraniata
Vertebrata/Craniata
Protochordata
Primitive Chordates
Exclusively marine
Vertebrae and paired appendages absent
On the basis of location of Notochord,divided in to four groups:-
Hemichordata
Calcichordata
Urochordata
Cephalochordata
7.
Hemichordata
Hemichordata(Acronworms)have beenshunted from Non-Chordata or Chordates and vice-versa
In fact they should be placed as a phylum of Non-Chordata
Every effort to homologize a peculiar forword extension of the gut in to the proboscis of Hemichordates wiyh the notochord
has failed to satisfy the modern zoologists
Characteristic Features of Phylum Hemichordata
Hemichordata is bilaterally symmetrical and triploblastic animal.
They are exclusively marine animals.
They can be solitary or in colonies.
Hemichordata have a true body cavity or coelom.
The digestive tract is complete with an anus and can be in the form of a U shaped tube or straight.
A buccal diverticulum is present in the proboscis.
Body is divided into three regions – Proboscis, Collar and Trunk.
Circulatory system in open type.
Respiration occurs through gills.
The proboscis has a glomerulus, which is the excretory organ.
A primitive nervous system is present.
Sexes are separate or united.
8.
Calcichordata
The primitivestem-chordates establishing a link between Echinoderms and
the Chordates
All are fossils
They had a skeleton of calcite plates,like Echinoderms
A notochord ran the length of the pleated tail
In a groove above the notochord,a nerve cord was also present
Gill slits were present
Example:- Cothurnocystis & Mitrocystella.
9.
Urochordata
“Oura” meansTail;Chorda means Cord
Notochord only present only in tail region of larva
Commonly called “Sea Squirt” or “Tunicates”
Exclusively marine ;solitary or colonial
Mostly sessile/fixed
Filter feeding
Nerve cord present in larva but replaced by a dorsal ganglion in adults
The pharyngeal slits are numerous ,persists in adults open in to an ectoderm lined cavity called
“Atrium”
There are no gills
Retrogressive metamorphosis
10.
Continued……………..
Body isgenerally covered or enclosed in a leathery bag like structure called
“Tunic” or “Test”
Tunic composed of tunicin (similar to cellulose)
These animals are also known as Tunicata
Larva is more developed as compare to adult
Larva called tadpole (undergo retrogressive metamorphosis)
The Ascidian largest class of tunicates
Examples:- Herdmania ,Ascidia,Pyrosoma,Oikopleura
11.
Cephalochordata
“Cephalos” meansHead ; Chorda means Cord
Members are commonly known as Lancelets
Only Cephalochordates having all four Chordate characters
Exclusively marine
Filter feeding
Notochord extends from head to tail region
Notochord persistent throughout their life but no brain formation
Numerous pharyngeal gill slits; persist in adults; open in to an Atrium instead
of exterior
Tail persists throughout life
12.
Continued………………
Small body(upto 5 cm ); elongated & laterally flattened
Nearly transparent tadpole creatures
Do not possess true heart ; major vessels pump blood
Dioecious; having numerous gonads
No asexual reproduction
Free swimming larva
Examples:- Branchiostoma(Amphioxus/Lancelets)
Present in shallow Sea water
Long trunk , short tail
Tail bear caudal fin
13.
Reference
Biology ofChordates
Dr. Harish C. Nigam
Modern Text Book of Zoology
Prof. R. L. Kotpal
www.biomentorsonline.com