Growth of coral reefs is affected by both abiotic and biotic factors. Abiotic factors that influence coral reef growth include latitude, temperature, light, salinity, turbidity, wave action, aerial exposure, and the need for a firm substrate. Biotic factors include mutualistic relationships between corals and zooxanthellae, as well as competition and predation between coral species. Both natural stresses such as storms, disease, and river runoff, as well as human impacts like pollution, sedimentation, and climate change threaten the growth and survival of coral reefs.