•WEL
COME…..
Submitted bySubmitted by
YAMGAR SAMBHAJI VISHNU.YAMGAR SAMBHAJI VISHNU.
Ph.D –I YEARPh.D –I YEAR
•PRESENTATION ON
Submitted toSubmitted to
Dr. N.S. Kute.Dr. N.S. Kute.
Seminar InchargeSeminar Incharge
Department of Agricultural BotanyDepartment of Agricultural Botany
(Genetics and Plant breeding).(Genetics and Plant breeding).
MAHATMA PHULE KRUSHI VIDYAPEETH, RAHURI.MAHATMA PHULE KRUSHI VIDYAPEETH, RAHURI.
HERITABILITY AND GENETIC ADVANCE
•2
• HeritabilityHeritability
•IntroductionIntroduction
•
• conclusionconclusion
•
Estimation of heritabilityEstimation of heritability
•Types of Heritability.Types of Heritability.
•Broad sence HeritabilityBroad sence Heritability
•Narrow sence heritability.Narrow sence heritability.
..
•3
 .
  
Heritability is the proportion of the total phenotypic
variation controlled by genetic rather than
environmental factors.
or
It is good index of transmission of character from parent
to their
Offspring (Falconer,1960).
Heritability H = VG / VP X 100
VP= VG+ VE
Where
The total phenotypic variance may be decomposed:
VP = total phenotypic variance
 VG = total genetic variance
 VE = environmental variance
 Broad sense heritability: (Lush,1940)
It is the ratio of genotypic variance (VG) to the total
phenotypic variance (VP)
h2
(bs) = VG / VP X 100
VP= VG+ VE
The total phenotypic variance may be
decomposed:
VP = total phenotypic variance
 VG = total genetic variance
 VE = environmental variance
 Heritability values are characterized as fallow.
Low- less than 30%
Moderate: 30-60%
High- More than 60% (Johnson et al. 1955).
 Useful in selection of elite types from homozygous
population.
 Can be estimated from both parental and segregating
material.
 Estimated from total genetic variance
 Narrow sense heritability:
It is the ratio of additive genetic variance (VA) to the total
phenotypic variance (VP) (Smith, 1952).
h2
(ns) = VA/ VP x 100
 . The total genetic variance (VG) may be decomposed:
 VA = additive genetic variance
 VD = dominance genetic variance
 VI = epistatic (interactive) genetic variance
Summary: Vp= VA+ VD+ VI+ VE+ VGE
 The magnitude of narrow sense heritability is always
less than or equal to broad since heritability.
 Useful in selection of elite types from segregating
material.
 Narrow sense heritability is classified as follow.
 Low-5 to 10 %
 Medium-10-30%
 High- more than 30% Robinson et.al(1966)
Broad sense heritability-
It require estimation of genetic variance in a population .
It is estimated as two way
1) Simple trials-h2
(bs) = VG / VP X 100
VP= VG+ VE
The total phenotypic variance may be decomposed:
VP = total phenotypic variance
VG = total genetic variance
VE = environmental variance
2) From Generation mean analysis-
( VF2 - VF1) / VF2 X 100
Where VF2, VF1 are variance of F1 and F2 respectively
For the estimation of narrow sense heritability require estimation of
additive genetic variance
1)Diallel Analysis
h2
(ns) = (1/2 D + ½ H1- ½ H2 – ½ F) X 100
(1/2 D + ½ H1- ¼ H2 – ½ F+ E)
2) Generation mean analysis.
Heritability= ½ D / VF2 X 100
Warner (1952)….
Heritability = D/ (D+H+E) X 100
Mather (1949)…
Broad sense heritability Narrow sense heritability
 Estimated from total
genetic variance.
 Can be estimated from
both parental and
segregating material.
 More useful in animal
breeding.
 Useful in selection of
elite types from
homozygous population.
 Estimated from additive
genetic variance.
 Requires crossing in a
definite fashion.
 Useful in both plant and
animal breeding.
 Useful in selection of
elite types from
segregating material.
 Genetics population
 Sample size- random Sampling.
 Method of computation-
Variance component method , Parent offspring
regression.
 Because it allows us to predict a trait’s
response to selection
R = h2
S
R = response to selection
h2
= heritability
S = selection differential


 It is useful in predicting the effectiveness of selection.
 It is also helpful for deciding breeding methods to be
followed, for effective selection.
 It gives an idea about the response of various
characters to selection pressure.
 It is useful in predicting the performance under
different degree of intensity of selection.
 It helps for construction of selection index.
 Estimates of heritability serve as a useful guide to the
breeder, to appreciate the proportion of variation,
which is due to genotypic or additive effects.
  
The Limitations of Heritability
• Heritability does not indicate the degree to which a characteristic is
genetically determined.
• Pure breed no polydactilly rabbits: still polydactilly can happen
• An individual does not have heritability.
• Narrow-sense heritability of 0.6 in population does not indicate that an individual’s
characteristic is 60% additive
• There is no universal heritability for a characteristic.
• Two populations will have different heritability due to environment
• Even when heritability is high, environmental factors may
influence a characteristic.
• Human height
• Heritability indicates nothing about the nature of population
differences in a characteristic.
 If broad scenes heritability is high character is
least influenced by Env. Selection is not desirable.
 If broad sense heritability is low character
highly influenced by Env.Due to masking effect of
genes.
 If narrow heritability is high character is governed
by additive fixable genes.(selection method)
 If narrow heritability is low character is governed
by non-additive ,non-fixable genes.
(Heterosis breeding Useful)
 Improvement in the mean genotypic value of
selected plants over the parental population is
known as genetic advance.
 Gs = (K) (σp) (H)
 Where, Gs: Genetic advance under selection
 K : Selection differential
σp : Phenotypic standard deviation of base population
 H : heritability
Genetic variability: Greater the variability,
higher will be the genetic advance and vice
versa.
Heritability: genetic advance is generally
high with the character having high
heritability and vice versa.
Selection intensity: It is the proportion of
plants selected from population. The high
selection intensity generally gives better
results than the low selection intensity.

Estimates of GA are based on empirical results and
free from genetic improvement under selection for
polygenic traits.
GA can be estimated both from parental as well as
segregating population.
Estimates of GA helps in understanding the types of
gene action involved into the expression of various
polygenic characters. High values of GA are
indicative of additive gene action.
It also helps in deciding a breeding procedure for the
genetic improvement of various polygenic traits by
determining the gene actions.

 Disadvantages of genetic advance:
 Estimates of GA are based on variance and
therefore, not statistically very robust.
 Estimates of GA are specific to the breeding
material and hence it varies according to the
breeding material.
If value of genetic value is high show character
is governed by additive gene (Selection
method rewarding)
If value of genetic value is low show character
is governed by non-additive genes(Heterosis
breeding rewarding)
•23
 High heritability with high genetic advance
character is governed by additive genes fixable
gene, Selection desirable.
 If heritability is high and low genetic advance
character is governed by non-additive ,fixable
genes.(selection not rewarding)
 If heritability is low with high genetic advance
character is governed by additive gene effects
 If heritability is low with low genetic advance
character is highly influenced by Env.
heritability its type and estimation of it

heritability its type and estimation of it

  • 1.
  • 2.
    Submitted bySubmitted by YAMGARSAMBHAJI VISHNU.YAMGAR SAMBHAJI VISHNU. Ph.D –I YEARPh.D –I YEAR •PRESENTATION ON Submitted toSubmitted to Dr. N.S. Kute.Dr. N.S. Kute. Seminar InchargeSeminar Incharge Department of Agricultural BotanyDepartment of Agricultural Botany (Genetics and Plant breeding).(Genetics and Plant breeding). MAHATMA PHULE KRUSHI VIDYAPEETH, RAHURI.MAHATMA PHULE KRUSHI VIDYAPEETH, RAHURI. HERITABILITY AND GENETIC ADVANCE •2
  • 3.
    • HeritabilityHeritability •IntroductionIntroduction • • conclusionconclusion • Estimationof heritabilityEstimation of heritability •Types of Heritability.Types of Heritability. •Broad sence HeritabilityBroad sence Heritability •Narrow sence heritability.Narrow sence heritability. .. •3
  • 4.
     .    Heritabilityis the proportion of the total phenotypic variation controlled by genetic rather than environmental factors.
  • 5.
    or It is goodindex of transmission of character from parent to their Offspring (Falconer,1960). Heritability H = VG / VP X 100 VP= VG+ VE Where The total phenotypic variance may be decomposed: VP = total phenotypic variance  VG = total genetic variance  VE = environmental variance
  • 6.
     Broad senseheritability: (Lush,1940) It is the ratio of genotypic variance (VG) to the total phenotypic variance (VP) h2 (bs) = VG / VP X 100 VP= VG+ VE The total phenotypic variance may be decomposed: VP = total phenotypic variance  VG = total genetic variance  VE = environmental variance
  • 7.
     Heritability valuesare characterized as fallow. Low- less than 30% Moderate: 30-60% High- More than 60% (Johnson et al. 1955).  Useful in selection of elite types from homozygous population.  Can be estimated from both parental and segregating material.  Estimated from total genetic variance
  • 8.
     Narrow senseheritability: It is the ratio of additive genetic variance (VA) to the total phenotypic variance (VP) (Smith, 1952). h2 (ns) = VA/ VP x 100  . The total genetic variance (VG) may be decomposed:  VA = additive genetic variance  VD = dominance genetic variance  VI = epistatic (interactive) genetic variance Summary: Vp= VA+ VD+ VI+ VE+ VGE
  • 9.
     The magnitudeof narrow sense heritability is always less than or equal to broad since heritability.  Useful in selection of elite types from segregating material.  Narrow sense heritability is classified as follow.  Low-5 to 10 %  Medium-10-30%  High- more than 30% Robinson et.al(1966)
  • 10.
    Broad sense heritability- Itrequire estimation of genetic variance in a population . It is estimated as two way 1) Simple trials-h2 (bs) = VG / VP X 100 VP= VG+ VE The total phenotypic variance may be decomposed: VP = total phenotypic variance VG = total genetic variance VE = environmental variance 2) From Generation mean analysis- ( VF2 - VF1) / VF2 X 100 Where VF2, VF1 are variance of F1 and F2 respectively
  • 11.
    For the estimationof narrow sense heritability require estimation of additive genetic variance 1)Diallel Analysis h2 (ns) = (1/2 D + ½ H1- ½ H2 – ½ F) X 100 (1/2 D + ½ H1- ¼ H2 – ½ F+ E) 2) Generation mean analysis. Heritability= ½ D / VF2 X 100 Warner (1952)…. Heritability = D/ (D+H+E) X 100 Mather (1949)…
  • 12.
    Broad sense heritabilityNarrow sense heritability  Estimated from total genetic variance.  Can be estimated from both parental and segregating material.  More useful in animal breeding.  Useful in selection of elite types from homozygous population.  Estimated from additive genetic variance.  Requires crossing in a definite fashion.  Useful in both plant and animal breeding.  Useful in selection of elite types from segregating material.
  • 13.
     Genetics population Sample size- random Sampling.  Method of computation- Variance component method , Parent offspring regression.
  • 14.
     Because itallows us to predict a trait’s response to selection R = h2 S R = response to selection h2 = heritability S = selection differential  
  • 15.
     It isuseful in predicting the effectiveness of selection.  It is also helpful for deciding breeding methods to be followed, for effective selection.  It gives an idea about the response of various characters to selection pressure.  It is useful in predicting the performance under different degree of intensity of selection.  It helps for construction of selection index.  Estimates of heritability serve as a useful guide to the breeder, to appreciate the proportion of variation, which is due to genotypic or additive effects.   
  • 16.
    The Limitations ofHeritability • Heritability does not indicate the degree to which a characteristic is genetically determined. • Pure breed no polydactilly rabbits: still polydactilly can happen • An individual does not have heritability. • Narrow-sense heritability of 0.6 in population does not indicate that an individual’s characteristic is 60% additive • There is no universal heritability for a characteristic. • Two populations will have different heritability due to environment • Even when heritability is high, environmental factors may influence a characteristic. • Human height • Heritability indicates nothing about the nature of population differences in a characteristic.
  • 17.
     If broadscenes heritability is high character is least influenced by Env. Selection is not desirable.  If broad sense heritability is low character highly influenced by Env.Due to masking effect of genes.  If narrow heritability is high character is governed by additive fixable genes.(selection method)  If narrow heritability is low character is governed by non-additive ,non-fixable genes. (Heterosis breeding Useful)
  • 18.
     Improvement inthe mean genotypic value of selected plants over the parental population is known as genetic advance.  Gs = (K) (σp) (H)  Where, Gs: Genetic advance under selection  K : Selection differential σp : Phenotypic standard deviation of base population  H : heritability
  • 19.
    Genetic variability: Greaterthe variability, higher will be the genetic advance and vice versa. Heritability: genetic advance is generally high with the character having high heritability and vice versa. Selection intensity: It is the proportion of plants selected from population. The high selection intensity generally gives better results than the low selection intensity. 
  • 20.
    Estimates of GAare based on empirical results and free from genetic improvement under selection for polygenic traits. GA can be estimated both from parental as well as segregating population. Estimates of GA helps in understanding the types of gene action involved into the expression of various polygenic characters. High values of GA are indicative of additive gene action. It also helps in deciding a breeding procedure for the genetic improvement of various polygenic traits by determining the gene actions. 
  • 21.
     Disadvantages ofgenetic advance:  Estimates of GA are based on variance and therefore, not statistically very robust.  Estimates of GA are specific to the breeding material and hence it varies according to the breeding material.
  • 22.
    If value ofgenetic value is high show character is governed by additive gene (Selection method rewarding) If value of genetic value is low show character is governed by non-additive genes(Heterosis breeding rewarding)
  • 23.
  • 24.
     High heritabilitywith high genetic advance character is governed by additive genes fixable gene, Selection desirable.  If heritability is high and low genetic advance character is governed by non-additive ,fixable genes.(selection not rewarding)  If heritability is low with high genetic advance character is governed by additive gene effects  If heritability is low with low genetic advance character is highly influenced by Env.