Evolution is the central theme of biology. It refers to change over time in allele frequencies in populations, driven primarily by natural selection. Speciation occurs through mechanisms like allopatric speciation. Charles Darwin developed the theory of evolution by natural selection, which proposes that heritable traits better suited to an organism's environment will increase the organism's fitness and chances of survival and reproduction, driving populations to change over generations. Phylogenies illustrate evolutionary relationships and history based on homologous traits in ancestors.