This document discusses tooth structure and evolution in mammals. It describes the four main tissues that make up teeth - enamel, dentine, cementum, and pulp cavity. Teeth are either rooted, with the opening to the pulp cavity constricting, or rootless, where the opening does not constrict. There are two theories on the evolution of teeth from single-cusped reptilian teeth to multi-cusped mammalian teeth - the concrescence theory involving fusion of teeth and the Cope-Osborn theory of additional cusps forming in front of and behind the original cusp. Specializations of cheek teeth include hypsodont, brachydont, bunodont, lophodont, and selenodont structures